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Lederberg and Tatum in 1946 conducted a classical experiment in bacterial genetics using *Escherichia coli* to demonstrate genetic recombination. They worked with two different mutant strains of *E. coli*, each deficient in specific nutritional requirements. One strain could not synthesize certain amino acids, while the other lacked the ability to produce different essential compounds. When grown separately on minimal medium, neither strain was _________ ___ ________ __________ _______ ___ _________ _______ ____ _______.
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