Discuss the role of enzyme topoisomerases in DNA replication.

Topoisomerases play a crucial role in DNA replication by managing the topological challenges that arise during the process. DNA replication involves unwinding of the double helix to expose the template strands for the synthesis of new complementary strands. However, this unwinding leads to the accumulation of torsional strain ahead of the replication fork, which, if left ____ _________ _____ _________ _______ ____ _______ ______ ___ _________.
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