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The Hamilton Rule, formulated by W.D. Hamilton, is a principle in evolutionary biology that explains how altruistic behavior can evolve through natural selection. It is expressed mathematically as rB > C, where r represents the genetic relatedness between the actor and the recipient, B is the reproductive benefit gained by the recipient, and C is ___ __________ ______ ____ _______ ________ ____ _______ _____ ________ ________.
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