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Endosperm shows considerable variation in its mode of development among angiosperms, and these variants are important for understanding seed biology and nutrition. Based on the pattern of nuclear division and cell wall formation, endosperm development is mainly classified into three variants: nuclear, cellular, and helobial endosperm. In the nuclear type, the primary endosperm nucleus undergoes repeated mitotic divisions without cytokinesis, resulting in a large multinucleate cell _________ _______ ______ _________ _______ ___ ________.
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