Describe the role of bryophytes as indicators.

Bryophytes, including mosses, liverworts, and hornworts, play a crucial role as indicators of environmental conditions due to their sensitivity to changes in their surroundings. These non-vascular plants are particularly useful in monitoring ecosystem health, as they lack the complex structures found in higher plants and depend directly on the surrounding _______ ________ ______ __________ ______ __________ ______ ___ _________ _________ ___ ____.
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