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Ion-exchange chromatography separates proteins based on their net surface charge at a particular pH. Proteins carry positive or negative charges depending on their amino acid composition and the surrounding pH. In cation-exchange chromatography, negatively charged resins bind positively charged proteins, whereas anion-exchange chromatography uses positively charged resins to bind negatively charged _________ _____ ___ _____ __________ ___ ____ _____.
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